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Here we pursue this problem in the Drosophila melanogaster ON motion vision circuit 4, 5, in which we record the membrane potentials of direction-selective T4 neurons and of their columnar input elements 6, 7 in response to visual and pharmacological stimuli in vivo. Nonlinear, multiplication-like operations carried out by individual nerve cells greatly enhance the computational power of a neural system 1, 2, 3, but our understanding of their biophysical implementation is scant.
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